In simple terms
A friendly intro before the formal notes — no formulas yet.
The Ecosystem's Economy
Ecosystems run on an energy budget, where energy flows through once and is lost, while essential nutrients are constantly recycled. Productivity measures how efficiently organisms convert energy into biomass.
Think of an ecosystem's energy flow like your personal finances. Your gross salary is the total energy captured (Gross Productivity). After you pay for your essential living costs like rent and food (Respiration), the money you have left to save or spend on growth is your net income (Net Productivity). You can't recycle the money you've spent, it's gone, just like energy is lost as heat.
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First, identify the source of energy (usually the sun) and trace its path through different trophic levels, remembering that energy is lost at each step.
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Next, distinguish between Gross Productivity (total energy assimilated) and Net Productivity (energy left for growth after respiration). Use the formulas NPP = GPP - R and NSP = GSP - R.
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Then, analyse nutrient cycles like carbon and nitrogen by identifying the main storages (sinks) and the processes that move nutrients between them (flows or fluxes).
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Finally, evaluate human impacts on these cycles, such as burning fossil fuels (carbon cycle) or using fertilisers (nitrogen cycle), and their consequences.
Explore the concept
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1 more simulation for this topic — run them in the Simulations section below
Simulations
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1 simulation
- PhETIB 2.2
Energy Forms and Changes
On the Systems screen, choose an energy source, a converter and a user, tick Energy Symbols and follow each chunk of energy through the chain.
Try this
- Open Systems, tick Energy Symbols, choose the Sun and the solar panel, and follow light energy becoming electrical energy.
- Swap the source to the cyclist, feed her, and watch chemical energy become mechanical and thermal energy.
- Count the chunks that reach the light bulb against the chunks that drift away as heat.
Look for Energy is never destroyed (first law), yet every transformation sheds heat (second law): the same reason only a fraction of energy reaches the next trophic level.
Simulation by PhET Interactive Simulations, University of Colorado Boulder · Licensed to MarkScheme (public licence CC BY-NC 4.0 since 2026-03-30)
Key formulas
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Full topic notes
Formal explanation with the rigour you need for the exam.
Energy Flow and the Laws of Thermodynamics
Energy in almost all ecosystems originates from the sun. This solar energy is captured by autotrophs (producers) and converted into chemical energy through photosynthesis. This energy is then transferred through the ecosystem as one organism consumes another. However, this flow is unidirectional and inefficient.
First Law of Thermodynamics: Energy can be transferred and transformed, but it cannot be created or destroyed. In ecosystems, this means solar energy is converted to chemical energy, which is then transferred between trophic levels.
Second Law of Thermodynamics: During any energy conversion, some energy is lost as heat, and the entropy (disorder) of the universe increases. This explains why energy transfers are inefficient (the '10% rule') and why food chains are limited in length. Energy is lost as heat during respiration at each trophic level.
Ecosystem Productivity
Productivity is the rate at which biomass is generated in an ecosystem. It is a key measure of an ecosystem's health and ability to support life. We distinguish between primary productivity (by producers) and secondary productivity (by consumers).
For Producers (Autotrophs):<br><b>Net Primary Productivity (NPP) = Gross Primary Productivity (GPP) - Respiration (R)</b><br><br>GPP is the total energy fixed by photosynthesis. R is the energy used by the producers for their own life processes. NPP is the energy available to herbivores.
For Consumers (Heterotrophs):<br><b>Net Secondary Productivity (NSP) = Gross Secondary Productivity (GSP) - Respiration (R)</b><br><br>GSP is the energy assimilated from food (). R is the energy used by the consumer for its life processes. NSP is the formation of new biomass (growth and reproduction).
Nutrient Cycling: Carbon and Nitrogen
Unlike energy, which flows through an ecosystem and is lost, matter is cycled. Biogeochemical cycles describe the pathways of essential elements like carbon and nitrogen through the biotic (living) and abiotic (non-living) components of an ecosystem. We model these using storages (where the nutrient is held) and flows (the processes that move it).
Carbon Cycle: Key storages include the atmosphere (), oceans, biomass, and fossil fuels. Key flows are photosynthesis (atmosphere to biomass), respiration (biomass to atmosphere), combustion (fossil fuels to atmosphere), and diffusion (atmosphere to ocean).
Nitrogen Cycle: The main storage is the atmosphere (). Key flows include nitrogen fixation (by bacteria or lightning, converting to ammonia), nitrification (ammonia to nitrates), assimilation (uptake by plants), denitrification (nitrates to ), and ammonification (decomposition).
Human Impact: Burning fossil fuels adds excess to the atmosphere, enhancing the greenhouse effect. The industrial production of fertilisers (Haber-Bosch process) has doubled the rate of nitrogen fixation, leading to eutrophication in aquatic systems.
Worked examples
See the formulas applied — reveal one step at a time, like the exam.
In a temperate forest ecosystem, the Gross Primary Productivity (GPP) was measured to be 8,400 . The plant community's respiratory losses (R) were found to be 4,600 . Calculate the Net Primary Productivity (NPP) of this ecosystem.
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<b>1. State the formula:</b><br>NPP = GPP - R [1 mark]<br><br><b>2. Substitute the values:</b><br>NPP = 8,400 - 4,600 [1 mark for correct substitution]<br><br><b>3. Calculate the final answer with units:</b><br>NPP = 3,800 [1 mark for correct answer and units]<br><br><b>Total: 3 marks</b>
A population of rabbits in a meadow consumes 25,000 of plant matter. They produce 16,000 of faeces (egestion). Their respiratory heat loss is 7,500 . a) Calculate the Gross Secondary Productivity (GSP). b) Calculate the Net Secondary Productivity (NSP).
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<b>a) Calculate GSP:</b><br><b>1. State the formula:</b> GSP = Food eaten - Faecal loss [1 mark]<br><b>2. Substitute values:</b> GSP = 25,000 - 16,000 <br><b>3. Final Answer:</b> GSP = 9,000 [1 mark]<br><br><b>b) Calculate NSP:</b><br><b>1. State the formula:</b> NSP = GSP - R [1 mark]<br><b>2. Substitute values:</b> NSP = 9,000 - 7,500 <br><b>3. Final Answer:</b> NSP = 1,500 [1 mark]<br><br><b>Total: 4 marks</b>
How it all connects
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Glossary
Key terms for this topic — skim now; the Check step will test them.
- Gross Primary Productivity (GPP)
The total amount of chemical energy produced by autotrophs in an ecosystem, typically through photosynthesis, per unit area per unit time. It's the total 'income' of energy before any is used. Units: or .
- Net Primary Productivity (NPP)
The energy remaining as biomass after primary producers have accounted for their own respiratory losses (R). This is the energy available to the next trophic level. Formula: .
- Gross Secondary Productivity (GSP)
The total energy or biomass assimilated by consumers from their food. It is calculated as the mass of food eaten minus the mass of faecal loss. Formula: .
- Net Secondary Productivity (NSP)
The energy remaining as biomass after consumers have accounted for their respiratory losses (R). This represents the growth and reproduction of the consumer. Formula: .
- Respiration (R)
The process where organisms convert organic matter into carbon dioxide and water, releasing energy for metabolic processes. This represents an energy loss from the ecosystem as heat.
- 10% Rule
A generalisation stating that only about 10% of the energy from one trophic level is transferred and incorporated into the biomass of the next trophic level. The other 90% is lost, primarily as heat during respiration, or is not consumed.
- can't most organisms use
The strong triple bond between the two nitrogen atoms in gas makes it very stable and unreactive. It must be 'fixed' (converted into ammonia, nitrates, or nitrites) by nitrogen-fixing bacteria or lightning before it can be used by plants.
Name it
Read the meaning, then pick which of this lesson’s terms it describes. Miss one and you see what your choice really means.
The energy remaining as biomass after primary producers have accounted for their own respiratory losses (R). This is the energy available to the next trophic level. Formula: .
Quick check
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Teach it back
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Revision flashcards
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Key takeaways
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First Law of Thermodynamics: Energy can be transferred and transformed, but it cannot be created or destroyed. In ecosystems, this means solar energy is converted to chemical energy, which is then transferred between trophic levels.
Second Law of Thermodynamics: During any energy conversion, some energy is lost as heat, and the entropy (disorder) of the universe increases. This explains why energy transfers are inefficient (the '10% rule') and why food chains are limited in length. Energy is lost as heat during respiration at each trophic level.
Practice — then mark it
The whole point: a real Cambridge question, marked mark-by-mark.
Test Your Knowledge on Energy Flow and Nutrient Cycles
Test Your Knowledge on Energy Flow and Nutrient Cycles
Extra simulations & links
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Frequently asked
Checkpoint
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